D. A. Kadimaliev Dr.Sc. (Biol.), Professor, Mordovian State University (Saransk). E-mail: cadimded@yandex.ru V. V. Shutova Ph.D. (Biol.), Associate Professor, Mordovian State University (Saransk). E-mail: vshutova@yandex.ru S.A. Ibragimova Ph.D. (Biol.), Associate Professor, Mordovian State University (Saransk). E-mail: ibragimova-s@yandex.ru G. V. Maksimov Dr.Sc. (Biol.), Professor, Moscow State University named after M.V. Lovonosov. E-mail: gmaksimov@mail.ru